Hello Zdiddy,

Thanks for your post! Assuming you're using the default Finish-to-Start dependencies, the dependent task's Start Date will always depend on the latest of the preceding task's End Dates. If you're using a different predecessor type or a combination of different predecessor types, then the important thing to know is that the Start (or Finish) of the dependent task will always depend on the latest of the preceding task's linked Start or Finish date. I've included a screenshot below. 

I hope this helps!

Isaac J

Smartsheet Support